Crepet, W. L. and G. D. Feldman. 1991. The Earliest remains of grasses in the fossil record. American Journal of Botany 78 (7): 1010-1014.

Abstract: New fossils provide the earliest unequivocal evidence of grasses. Spikelets and inflorescence fragments with included pollen from the Paleocene/Eocene Wilcox Formation in western Tennessee have a suite of diagnostic characters that limits their affinities to Poaceae. Associated vegetative remains are also suggestive of grasses, but are not well enough preserved for an unequivocal identification. These fossils indicate a minimal time as wind pollinated Hamamelididae , wind pollination evolved in grasses or persisted in the grass lineage (if a wind-pollinated grass sister group is presumed) in dry tropical environments.


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Actual width in ( ) unless otherwise noted.

Rhizome with leaf sheaths
(image= 4 cm)
Spikelet with two florets
(height= 6 mm)
Pollen grain
(16 ┬Ám)